Significant Shareholders

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Shares in Issue: 530,412,678 1p Ordinary Shares

Last updated: 24th September 2020

Significant Shareholders	Amount	% Holding
Fidelity Management & Research	49,810,760	9.39
Slater Investment	42,301,894	7.98
John Dawson (beneficial and non-beneficial holdings)	44,212,402	7.30
BlackRock Inc	27,451,889	5.18
Kempen Capital Management N.V.	26,621,969	5.01
MVM Life Science Partners LLP	24,760,663	4.70
Polar Capital Holdings	19,663,707	3.71
Investec Group	18,187,402	3.43
GVQ Investment	18,006,489	3.39
Rathbone Investment	17,423,951	3.29

Shareholdings of Directors & their families	Amount	% Holding
Peter Butterfield	412,461	0.08
David Cook	234,129	0.04
Nigel Clifford	180,663	0.04
Andrew Franklin	128,384	0.02
Richard Jones	15,000	0.00