Significant Shareholders

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Shares in Issue: 519,490,041 1p Ordinary Shares

Last updated: 18 September 2019

Significant Shareholders	Amount	% Holding
John Dawson (beneficial and non-beneficial holdings)	55,476,402	10.68
Fidelity Management & Research	44,851,631	8.63
Slater Investment	41,446,094	7.98
BlackRock Inc	38,638,235	7.44
MVM Life Science Partners LLP	37,544,892	7.23
RIT Capital Partners Plc	23,397,024	4.50
Polar Capital Holdings	20,698,583	3.98
Investec Group	16,402,379	3.16

Shareholdings of Directors & their families	Amount	% Holding
Nigel Clifford	180,663	0.04
David Cook	102,371	0.02
Peter Butterfield	28,376	0.01
Richard Jones	15,000	0.00