Significant Shareholders

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Shares in Issue: 529,489,021 1p Ordinary Shares

Last updated: 4th February 2020

Significant Shareholders	Amount	% Holding
John Dawson (beneficial and non-beneficial holdings)	50,976,402	9.6
Fidelity Management & Research	48,447,412	9.19
Slater Investment	41,446,094	7.98
BlackRock Inc	38,638,235	7.44
MVM Life Science Partners LLP	30,535,598	5.77
RIT Capital Partners Plc	23,397,024	4.50
Polar Capital Holdings	20,698,583	3.98
Investec Group	16,402,379	3.16

Shareholdings of Directors & their families	Amount	% Holding
Nigel Clifford	180,663	0.04
Andrew Franklin	128,384	0.02
David Cook	102,371	0.02
Peter Butterfield	28,376	0.01
Richard Jones	15,000	0.00