Significant Shareholders

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Shares in Issue: 530,412,678 1p Ordinary Shares

Last updated: 24th November 2020

Significant Shareholders	Amount	% Holding
Fidelity Management & Research	50,465,122	9.48
Slater Investment	41,741,894	7.84
BlackRock, Inc	36,196,710	6.80
John Dawson (beneficial and non-beneficial holdings)	29,976,402	5.60
F Van Lanschot Bankiers	29,514,372	5.54
MVM Life Science Partners LLP	23,967,019	4.50
Polar Capital Holdings	19,696,984	3.70
Investec Group	19,475,062	3.66
Rathbone Investment	17,880,512	3.36

Shareholdings of Directors & their families	Amount	% Holding
Peter Butterfield	412,461	0.08
David Cook	234,129	0.04
Nigel Clifford	180,663	0.04
Andrew Franklin	128,384	0.02
Richard Jones	15,000	0.00