Significant Shareholders

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Shares in Issue: 530,412,678 1p Ordinary Shares

Last updated: 13th May 2020

Significant Shareholders	Amount	% Holding
Fidelity Management & Research	49,810,760	9.39
John Dawson (beneficial and non-beneficial holdings)	49,362,402	9.31
Slater Investment	42,301,894	7.98
BlackRock Inc	27,451,889	5.18
MVM Life Science Partners LLP	26,426,561	4.98
Polar Capital Holdings	19,663,707	3.71
Investec Group	18,187,402	3.43
GVQ Investment	18,006,489	3.39
Rathbone Investment	17,423,951	3.29
Artemis Investment	17179066	3.24

Shareholdings of Directors & their families	Amount	% Holding
Peter Butterfield	412,461	0.08
Nigel Clifford	180,663	0.04
Andrew Franklin	128,384	0.02
David Cook	102,371	0.02
Richard Jones	15,000	0.00