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Significant Shareholders

Shares in Issue: 530,412,678 1p Ordinary Shares

Last updated: 13th May 2020

Significant Shareholders

Amount

% Holding

Fidelity Management & Research

49,810,760

9.39

John Dawson (beneficial and non-beneficial holdings)

49,362,402

9.31

Slater Investment

42,301,894

7.98

BlackRock Inc

27,451,889

5.18

MVM Life Science Partners LLP

26,426,561

4.98

Polar Capital Holdings

19,663,707

3.71

Investec Group18,187,402

3.43

GVQ Investment

18,006,489

3.39

Rathbone Investment17,423,951

3.29

Artemis Investment17179066

3.24

Shareholdings of Directors & their families

Amount

% Holding

Peter Butterfield

412,461

0.08

Nigel Clifford

180,663

0.04

Andrew Franklin

128,384

0.02

David Cook

102,371

0.02

Richard Jones

15,000

0.00