Significant Shareholders

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Shares in Issue: 530,412,678 1p Ordinary Shares

Last updated: 24th November 2020

Significant Shareholders	Amount	% Holding
Fidelity Management & Research	53,317,753	9.98
Slater Investment	42,019,007	7.87
BlackRock, Inc	36,056,498	6.75
F Van Lanschot Bankiers	32,678,985	6.12
John Dawson (beneficial and non-beneficial holdings)	25,456,917	4.77
Polar Capital Holdings	22,207,692	4.16
Investec Group	20,687,183	3.87
Rathbone plc	18,428,800	3.45
Affiliated Managers Group	16,304,802	3.05
MVM Life Science Partners LLP	14,870,613	2.78

Shareholdings of Directors & their families	Amount	% Holding
Peter Butterfield	412,461	0.08
David Cook	234,129	0.04
Nigel Clifford	180,663	0.04
Andrew Franklin	128,384	0.02
Richard Jones	15,000	0.00