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Significant Shareholders

Shares in Issue: 519,490,041 1p Ordinary Shares

Last updated: 18 September 2019

Significant Shareholders

Amount

% Holding

John Dawson (beneficial and non-beneficial holdings)

55,476,402

10.68

Fidelity Management & Research

44,851,631

8.63

Slater Investment

41,446,094

7.98

BlackRock Inc

38,638,235

7.44

MVM Life Science Partners LLP

37,544,892

7.23

RIT Capital Partners Plc

23,397,024

4.50

Polar Capital Holdings

20,698,583

3.98

Investec Group

16,402,379

3.16

Shareholdings of Directors & their families

Amount

% Holding

Nigel Clifford

180,663

0.04

David Cook

102,371

0.02

Peter Butterfield

28,376

0.01

Richard Jones

15,000

0.00